Mantel tests (Mantel, 1967) and analyses of spatial autocorrelation were performed for both nuclear and mitochondrial markers as implemented in Alleles in Space (AIS) v1.0 (Miller, 2005) to determine the correlation between genetic and geographical distances across the study area. The spatial autocorrelation analyses were conducted using 10, 20 and 50 geographic distance classes of equal size. The significance of both tests was evaluated through the use of a randomisation procedure based on 1000 permutation replicates.