The remaining species of Echinocereus form a wellsupported clade (Echinocereus s. s.) that is sister to the Stenocereus group (Figs. 2, 3; 100% MP BS / 100% MP JK, 1.00 BI PP). Two main groups are recovered within the Echinocereus s. s. Group A has 11 taxa with weak to moderate support (