wwhere D is the detrital biomass in (g C m−2), PP is the primary pro-duction in (g C m−2); E is the euphotic depth in meters. The depthof the euphotic zone was calculated as follows: E = 2.5 × SD (Sec-chi depth in meters), the average SD of TSL is 1.07 m and thereforeE = 2.68 m.