Although morphological evidence for myriapod monophyly is often described as elusive, the details of the mobile cephalic endoskeleton (‘‘swinging tentorium’’) and its role in abduction of the mandibular gnathal lobe are unique to Myriapoda (Koch, 2003), and may be supplemented by arrangements of serotonin reactive neurons (Harzsch, 2004) and the position of the nuclei of eucone cells in the ommatidia (Mu¨ller et al., 2007) and the cuticle of intermediary cells in epidermal glandular organs (Hilken et al., 2005) that are shared by chilopods and diplopods and apparently apomorphic for Myriapoda.