In some plant and animal species, individuals can be cloned and their genetically identical copies can be tested in multiple environments. Cloning individuals for within family-selection is an especially common practice to develop varieties for deployment in eucalyptus species (Mullin et al., 2011) and many ornamental plants and vegetatively-propagated crops. Cloning allows breeders to capture hybrid vigor and non-additive genetic effects among and within crosses. Cloning individuals and testing in replicated field trials has the advantage of increasing the accuracy of genotypic value predictions compared to individual evaluations (Isik et al., 2004). With cloned progeny we can increase the heritability both by increasing the numerator (including all of the genotypic variance because broad-sense heritability is applicable to gain from
selection) and by decreasing the denominator by averaging clonal means over more observations. If progenies from a mating design are cloned, we can also estimate total genotypic variance in addition to additive variance, providing a means to assess the relative importance of non-additive genetic variance.