Where 兪 = overall mean, ..! = effect of location i, ..(..)!" = effect of block j nested within
location i (replication effect), ..! = effect of genotype k (RIL effect), ....!" = effect of interaction
between genotype k and location i, ..!"# = residual (experimental error) effect of the plot
containing genotype k in block j of location i. We will assume that all effects except the overall
mean are random.